Main scientific papers

  • Geographic variation in morphometrics, molt, and migration suggests ongoing subspeciation in Pacific Golden-Plovers (Pluvialis fulva). Auk 132 (2015): 647-656 (Joop Jukema, Johan van Rhijn & Theunis Piersma).
  • Diversity of nuptial plumages in male Ruffs Philomachus pugnax. Ardea 102 (2014): 5-20 (Johan van Rhijn, Joop Jukema & Theunis Piersma).
  • In tundra plovers the frequency of inner flight feather replacement varies with length of long-distance flights. Ardea 101 (2013): 121-132 (Joop Jukema, Johan van Rhijn, Peter Olsson & Theunis Piersma).
  • Incomplete and irregular annual replacement of secondaries in Eurasian Golden Plovers, Pluvialis apricaria. Wader Study Group Bull. 120 (2013): 102-107 (Joop Jukema, Erik Bunskoeke, Theunis Piersma, Ton Pieters, Anita Koolhaas & Johan van Rhijn).
  • Mate guarding as a key factor in the evolution of parental care in birds. Anim.Behav. 41 (1991): 963-970.
  • Unidirectionality in the phylogeny of social organisation, with special reference to birds. Behaviour 115 (1990): 153-174.
  • On the mechanism of mate selection in black-headed gulls. Behaviour 100 (1987): 134-169 (Johan van Rhijn & Ton Groothuis).
  • Biparental care and the basis for alternative bond-types among gulls, with special reference to Black-headed Gulls. Ardea 73 (1985b): 159-174 (Johan van Rhijn & Ton Groothuis).
  • Black-headed gull or black-headed girl? On the advantage of concealing sex in gulls and other colonial birds. Neth.J. Zool. 35 (1985): 87-102.
  • A scenario for the evolution of social organization in Ruffs Philomachus pugnax and other charadriiform species. Ardea 73 (1985a): 25-37.
  • Phylogenetical constraints in the evolution of parental care strategies in birds. Neth.J.Zool. 34 (1984): 103-122.
  • On the maintenance and origin of alternative strategies in the Ruff Philomachus pugnax. Ibis 125 (1983): 482-498.
  • Units of behaviour in the black-headed gull, Larus ridibundus L. Anim.Behav. 29 (1981): 586-593.
  • Being honest about one's intentions: an Evolutionary Stable Strategy for animal conflicts. J.theor.Biol. 85 (1980): 623-641 (Johan van Rhijn & Ron Vodegel).
  • Communication by agonistic displays: a discussion. Behaviour 74 (1980): 284-293.
  • Processes in feathers caused by bathing in water. Ardea 65 (1977): 126-147.
  • The patterning of preening and other comfort behaviour in a herring gull. Behaviour 63 (1977): 71-109.
  • Behavioural dimorphism in male ruffs. Behaviour 47 (1973): 153-229.
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